Background
Streptococcus mitis is a major cause of infective endocarditis (IE), a life-threatening infection of the heart valves. S. mitis may harbour capsule biosynthesis loci (cps), a key virulence factor for Streptococcus pneumoniae (the pneumococcus). However, the prevalence and diversity of S. mitis strains carrying pneumococcal-like capsules remain poorly characterised. We investigated the occurrence of invasive S. mitis strains with pneumococcal-like serotype capsules.
Methods
We conducted retrospective whole-genome sequencing of 129 S. mitis isolates collected from patients with bloodstream infections and clinically diagnosed IE in the UK and Ireland from 2001-2016. Isolates were obtained from the British Society of Antimicrobial Chemotherapy and the UK Health Security Agency. Population structure and cps were analysed using comparative genomics and novel in silico tools.
Results
We identified 127/129 (98.4%) unique multilocus sequence types and Global Sequence Clusters (GSCs) among the isolates. Complete cps loci were identified in 116/129 (89.9%) isolates, with sequence lengths ranging from 6,628-32,574bp (median: 21,782bp). Pneumococcal-like vaccine serotypes 5 and multiple non-vaccine serotypes including 19C and 23B were detected among 26/116 (22.4%) cps using the pneumococcal SeroCall tool, with nucleotide-BLAST coverage and identity ranging from 82–100% and 86–95.76%, respectively. Novel cps loci were identified in 90/116 (77.6%) of the isolates.
Conclusion
The genomic overlap between S. mitis and pneumococcal cps loci highlights the close evolutionary relationship between the species. Extensive S. mitis cps diversity has the potential to drive pneumococcal vaccine escape facilitated by interspecies horizontal gene transfer at the cps locus.