Background
The MORDOR azithromycin mass drug administration (MDA) cluster-randomised trial conducted in sub-Saharan Africa (2014–2018) demonstrated a 13.5% reduction in mortality in communities receiving biannual azithromycin versus placebo. However, the impact of MDA on the dissemination of macrolide resistance via mobile integrative conjugative elements (ICEs) in Streptococcus pneumoniae remains poorly understood. We investigated potential horizontal gene transfer (HGT) of ICEs among pneumococci across a previous MDA trial site.
Methods
We performed longitudinal, geographically weighted sampling of children aged 1–9 years in the Malawi MORDOR trial site at baseline (2015), 6-months post-MDA (2017), and 3.5-years post-MDA (2021). Whole-genome sequencing was conducted on 495 prospective and 392 historical pneumococcal isolates from trial clusters. Global pneumococcal sequence clusters (GPSCs), sequence types (STs), antimicrobial resistance (AMR) genes, ICEs and their sequence diversity were analysed.
Results
Complete ICEs were identified in 53% (169/318) of macrolide-resistant isolates using ICEFinder. The most prevalent ICE family was Tn2010 (~20 kb), which carries macrolide resistance genes mefA and tetracycline resistance gene tet, and was detected in 135 isolates. Despite extensive Tn2010 diversity, several sequences clustered by similarity and identical ICEs were found in pneumococcal isolates from distinct genetic backgrounds, including vaccine-type (VT) serotype 6A (GPSC180-ST9549) and non-vaccine-type (NVT) serotype 15A (GPSC5-ST172).
Conclusion
The presence of identical ICEs in strains from diverse genetic backgrounds and serotypes suggests HGT, which could limit antibiotic options in disease caused by NVTs acquiring these elements. High-resolution genomic surveillance is essential to track ICE movement across lineages to inform future vaccine interventions.