Background
Encounters with the ubiquitous human commensal and common pathogen Streptococcus pyogenes occur repeatedly over the human lifespan. There are still critical areas of uncertainty regarding how the human immune system recognises, responds to, and remembers S. pyogenes, hindering modern vaccine development. Here, we investigated B cell responses to leading S. pyogenes candidate vaccine antigens.
Methods
The recently developed human challenge model of emm75 S. pyogenes pharyngitis in healthy adults (CHIVAS-M75) presents a unique opportunity to characterise pre-existing immunity and the nature of recalled immune memory upon repeat exposure. Using peripheral blood mononuclear cells collected before and after challenge, we used flow cytometry and single-cell sequencing approaches to compare B cell responses to S. pyogenes vaccine antigens (including SpyCEP, SpyAD, SCPA, and SLO) in CHIVAS-M75 challenge trial participants, comparing those who did and did not develop pharyngitis.
Results
We characterised how infection changes the magnitude, gene expression, phenotype, and B cell receptor repertoires of vaccine antigen-specific memory B cells. Overall, pre-existing memory B cell responses to were common in these healthy adults, and responses were modified by the experimental infection. The proportion of antigen-specific isotype class-switched B cells increased post-challenge, suggesting a maturation of cellular memory upon pathogen encounter. Increases in antigen-specific B cells correlated with antigen-specific serum IgG at 3 months post-challenge.
Conclusion
Our work provides foundational insights into how a single infection shapes immunity and its relationship to symptomatic infection, and suggests that pre-existing B cell memory may influence S. pyogenes vaccine immunogenicity in clinical trials in adults.